鞘蕊属
（重定向自Coleus）
| 鞘蕊花属File:Infobox info icon2.svg | |
|---|---|
| File:Leaf -pani koorkka.JPG | |
| 左手香（Coleus amboinicus；syn. Plectranthus amboinicus） | |
| 科学分类 编辑 | |
| 界： | 植物界 Plantae |
| 演化支： | 维管植物 Tracheophyta |
| 演化支： | 被子植物 Angiosperms |
| 演化支： | 真双子叶植物 Eudicots |
| 演化支： | 菊类植物 Asterids |
| 目： | 唇形目 Lamiales |
| 科： | 唇形科 Lamiaceae |
| 亚科： | 荆芥亚科 Nepetoideae |
| 族： | 罗勒族 Ocimeae |
| 属： | 鞘蕊花属 Coleus Lour.[1] |
| 异名[1] | |
| |
鞘蕊属（学名：Coleus），或作鞘蕊花属，是一个一年生或多年生的草本或灌木植物的属。部分物种属于多肉植物，长有具肉质或块状的块茎或块根。 分布于旧世界（东半球）的热带与亚热带。 有关鞘蕊属、彩叶草属及马刺花属三者的关系很混乱：前两属的物种曾经被并归马刺花属，但结果使马刺花属在支序亲缘分析之下变成了马刺花亚族（Plectranthinae）的一个多系群。
现时最新的分类学处理重新建立鞘蕊属，212个物种分别来自马刺花属、密穗花属及排香草属；Equilabium在系统发育研究支持下从马刺花属独立出去成为一个属[2]。
鞘蕊属物种常被用作园艺植物而种植，当中尤以彩叶草最为常见。一方面，彩叶草的斑斓叶片使之成为一种受欢迎的园艺植物，而近年兴起的生酮饮食，亦因为彩叶草的根部丰富的营养素而使之大受欢迎。彩叶草喜光，但不适宜太阳直射，而是非直射的光线：因为直射的太阳光会破坏叶片上的色素，使叶片颜色变得单一；所以在户外时，彩叶草应完全遮光，只能照射日出时最柔和的日光[3]。
其他长有块根的鞘蕊属植物有被当作食粮而种植，计有Coleus esculentus、Coleus rotundifolius及Coleus maculosus subsp. edulis。
分类学[编辑]
鞘蕊属最早由若昂·德理路于1790年描述。以形态学特征为指导基础，将它由马刺花属（首先由夏尔·莱里捷在1788年描述）独立出来；其四条雄蕊融合在一起，而不是从基底起就已分开。1962年，约翰·肯尼斯·莫顿留意到雄蕊融合在一起的物种比过往所认知的其实更常见及广泛分布，所以把鞘蕊属物种合并进马刺花属，而彩叶草属（Solenostemon）及一些其他的属仍维持区隔。[4]大多数的来源都跟随莫顿进行鞘蕊属的合并；有些保持彩叶草属的独立，但其他则将它跟着鞘蕊属一起并入马刺花属。[2]
在2004年针对唇形科荆芥亚科罗勒族（Ocimeae）进行的初步研究表明，马刺花亚族（Plectranthinae）为单系群，有两个主要演化支：一个包含鞘蕊属的模式种，并包含彩叶草属；另一个包含马刺花属的模式种以及其他属，因此广义上马刺花属不是单系群。在2018年进行的更详细研究得出类似的结论，并建议应重新确认鞘蕊属（包含彩叶草属），对马刺花属进行更狭义的定义，并建立一个新的属Equilabium，主要来自热带非洲的前马刺花属物种演化支。2019年提供了实现此方法所需的许多新的双名组合。[2]
支序亲缘[编辑]
裴顿等人(2019)[2]基于其2018年的研究总结[5]，出版了马刺花亚族（Plectranthinae）各属的支序亲缘图。下图中被裴顿等人接纳并构成广义的马刺花属的三个属以黄色底显示：
| 罗勒族 Ocimeae |
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物种[编辑]
Paton et al. (2019)列出本属物种有294种，当中大多数均由马刺花属转移过来[2]。截至2020年7月[update]，《世界植物在线》列出本属25个物种[1]。部分物详列如下[2]：
- Calchas parvifolius
- Coleus abyssinicus Coleus albicalyx
- Coleus betonicifolius
- Coleus linearifolius (J.K.Morton) A.J.Paton Coleus cuneatus
- (Baker) A.J.Paton Baker f.
- A.J.Paton
- Coleus longipetiolatus Coleus angolensis
- Coleus cyanophyllus Coleus longipetiolatus
- (P.I.Forst.) P.I.Forst. Gürke ex Engl., 1894 Coleus bifidus
- Coleus luengerensis Gürke
- Coleus cylindraceus (Suddee) Suddee (G.Taylor) A.J.Paton
- (A.J.Paton) A.J.Paton (Hochst. ex Benth.) A.J.Paton (A.Chev.) Roberty
- Coleus macrostachys
- Coleus daviesii
- Coleus maculosus E.A.Bruce (Lam.) A.J.Paton
- Coleus bipartitus
- Coleus madagascariensis ，异名：Plectranthus madagascariensis）又名：马达加斯加延命草、马达加斯加香茶菜 Coleus decimus
- (P.I.Forst.) P.I.Forst. (A.J.Paton) A.J.Paton
- P.I.Forst. & A.J.Paton Coleus actites
- Coleus angulatus
- Coleus decurrens Coleus megacalyx
- Gürke (A.J.Paton) A.J.Paton Coleus bishopianus
- Coleus megadontus (P.I.Forst.) P.I.Forst.
- Coleus deflexifolius (Hedge) A.J.Paton & Phillipson
- (Gamble) Smitha & A.J.Paton (Baker) A.J.Paton (Baker) A.J.Paton & Phillipson
- Coleus meyeri
- Coleus defoliatus
- Coleus minor (Hochst. ex Benth.) A.J.Paton Coleus aliciae
- Coleus blakei
- Coleus minutiflorus (Ryding) A.J.Paton Coleus densus
- (P.I.Forst.) P.I.Forst. (N.E.Br.) A.J.Paton
- (P.I.Forst.) P.I.Forst.
- Coleus mirus Coleus apoensis
- Coleus descampsii Coleus mitis
- (Briq.) A.J.Paton (R.A.Clement) A.J.Paton Coleus bojeri
- Coleus monostachyus (Fresen.) A.J.Paton
- (P.I.Forst.) P.I.Forst. (Codd) A.J.Paton Elmer
- Benth. (Robyns & Lebrun) A.J.Paton Hochst. ex Ces.
- Coleus mutabilis
- Coleus dichotomus
- Coleus myrianthellus (A.J.Paton) A.J.Paton Briq.
- Coleus bolavenensis
- Coleus namuliensis E.Downes & I.Darbysh. Coleus dinteri
- [[Suddee, Tagane & Rueangr.[6]]] (Briq.) A.J.Paton
- (Schltr.) Codd
- Coleus apreptus
- Coleus divaricatus Coleus nervosus
- A.J.Paton Briq. Coleus botryosus
- Coleus niamniamensis (Gürke) A.J.Paton
- Coleus diversus (S.T.Blake) P.I.Forst. & T.C.Wilson
- (A.J.Paton) A.J.Paton (S.T.Blake) P.I.Forst. & T.C.Wilson (Alston) A.J.Paton
- Coleus nitidus
- Coleus dolichopodus
- Coleus niveus Coleus alloplectus
- Coleus bourneae
- Coleus omahekense Dinter Coleus dumicola
- (Gamble) Smitha & A.J.Paton (P.I.Forst.) P.I.Forst.
- (P.I.Forst.) P.I.Forst.
- Coleus orbicularis Coleus apricus 光萼鞘蕊花
- Coleus dysophylloides Coleus orthodontus
- (Benth.) A.J.Paton (Gürke) A.J.Paton Coleus bracteatus
- Coleus otostegioides (Schweinf. ex Gürke) A.J.Paton
- Coleus elliotii (S.T.Blake) P.I.Forst. & T.C.Wilson (P.I.Forst.) P.I.Forst.
- Dunn (S.Moore) A.J.Paton (Wall. ex Benth.) A.J.Paton
- Coleus parensis
- Coleus eminii
- Coleus parishii (Gürke) A.J.Paton (Hook.f.) A.J.Paton
- Coleus brazzavillensis
- Coleus parvicalyx (A.J.Paton) A.J.Paton Coleus engleri
- A.Chev. (Briq.) A.J.Paton
- Benth. Calchas platystomoides
- Coleus adenanthus Coleus arabicus
- Coleus erici-rosenii
- Coleus penzigii (R.E.Fr.) A.J.Paton Schweinf. ex Penz.
- Coleus buchananii 食用鞘蕊花 Coleus perrieri
- (Hedge) A.J.Paton & Phillipson Coleus esculentus
- Benth. (Baker) Brenan ，异名：Plectranthus esculentus）
- Briq.
- Coleus petraeus Coleus eungellaensis
- Coleus petricola A.J.Paton
- Coleus alpinus Coleus burorum
- Coleus petrophilus Gürke
- Coleus excelsus Chiov.
- (P.I.Forst.) P.I.Forst. (Suddee, Suphuntee & Saengrit) Suddee
- Coleus plantagineus Coleus arenicola
- Coleus exilis
- Coleus platyphyllus (A.J.Paton) A.J.Paton (A.J.Paton) A.J.Paton
- Coleus caillei Coleus polystachyus
- (Benth.) A.J.Paton (Dalzell & A.Gibson) A.J.Paton Vatke
- (P.I.Forst.) P.I.Forst. A.Chev. (P.I.Forst.) P.I.Forst.
- (van Jaarsv. & P.J.D.Winter) A.J.Paton
- Coleus porphyranthus Coleus ferricola
- Coleus prittwitzii Phillipson, O.Hooper & A.J.Paton
- (Perkins) A.J.Paton 银叶到手香 Coleus caillei
- Coleus prostratus (Gürke) A.J.Paton
- Coleus foetidus A.Chev. ex Hutch. & Dalziel
- (Benth.) A.J.Paton (Codd) A.J.Paton
- Coleus argentatus
- Coleus foliatus
- Coleus pulchellus (A.J.Paton) A.J.Paton (P.I.Forst.) P.I.Forst.
- Coleus calaminthoides 兰屿小鞘蕊花 Coleus pumilius
- Blanco, 1837 Coleus formosanus
- (S.T.Blake) P.I.Forst. & T.C.Wilson；又名：银叶马刺花、银叶香茶菜，异名：''Plectranthus argentatus'' (Baker) A.J.Paton
- Kiew, 2021
- Coleus recurvata Coleus forsteri
- Coleus acariformis
- Coleus altanmouiensis Coleus calcicola
- Coleus robustus (Hook.f.) A.J.Paton
- Coleus fragrantissimus 圆叶鞘蕊花 Murata
- (P.I.Forst.) P.I.Forst. (Poir.) M.R.Almeida, 2003，异名：Plectranthus rotundifolius
- Coleus ruandensis Coleus argenteus
- Coleus fredericii
- Coleus rutenbergianus ，异名：Plectranthus welwitschii (Vatke) A.J.Paton & Phillipson
- Coleus caldericola Coleus sahyadricus
- (Smitha & Sunojk.) Smitha Coleus gamblei (T.C.Wilson, P.I.Forst. & M.A.M.Renner) T.C.Wilson & P.I.Forst.
- (Gamble) A.J.Paton (P.I.Forst.) P.I.Forst. (Smitha & Sunojk.) Smitha
- (A.J.Paton) A.J.Paton
- Coleus sanguineus Coleus geminatus
- Coleus saxicola (P.I.Forst.) P.I.Forst.
- Gürke Coleus calycinus
- Coleus saxorum (J.Mathew, Yohannan & B.J.Conn) Smitha
- Coleus gibbosus (Benth.) A.J.Paton
- A.J.Paton (Benth.) A.J.Paton
- Coleus adenophorus 到手香 Coleus argentifolius
- Coleus gigantifolius
- Coleus schliebenii (Suddee) Suddee (Mildbr.) A.J.Paton
- Coleus cambodianus Coleus scruposus
- A.J.Paton Coleus gillettii 彩叶草
- (Ryding) A.J.Paton (Murata) A.J.Paton (J.K.Morton) A.J.Paton
- (L.) Benth.，异名''Coleus blumei'', ''Plectranthus scutellarioides''
- Coleus scutellarioides Coleus glabriflorus
- Coleus sessilifolius (P.I.Forst.) P.I.Forst.
- Coleus amboinicus Coleus caninus
- Coleus shoolamudianus (Sunil & Naveen Kum.) Smitha & A.J.Paton
- Coleus globosus (Roth) Vatke，异名：Plectranthus caninus
- (Ryding) A.J.Paton (A.J.Paton) A.J.Paton
- Coleus socotranus Coleus aromatium
- Coleus goetzenii
- Coleus sphaerocephalus (Gürke) A.J.Paton (Baker) A.J.Paton
- Coleus caninus (Benth.) P.V.Heath, 1999
- (Robyns & Lebrun) P.V.Heath, 1999 (P.I.Forst.) P.I.Forst. (Gürke) A.J.Paton
- [[Lour.，异名：Plectranthus amboinicus）又名：左手香、过手香]] Benth. Paton, Mwanyambo, Govaerts, Smitha, Suddee, Phillipson, Wilson, Forster & Culham, 2019
- A.J.Paton A.Chev.
- Coleus steenisii 肉叶鞘蕊花
- Coleus gracilipedicellatum Coleus stenostachys
- (Robyns & Lebrun) A.J.Paton (Baker) A.J.Paton & Phillipson
- Coleus carnosifolius Coleus strictipes
- (G.Taylor) A.J.Paton Coleus graminifolius
- (Hemsl.) Dunn (Perkins) A.J.Paton (Roxb.) A.J.Paton
- Coleus articulatus Coleus grandidentatus
- Coleus suaveolens (Gürke) A.J.Paton
- (S.T.Blake) P.I.Forst. & T.C.Wilson Coleus casamancicus
- Coleus subscandens Gürke Coleus graniticola
- (I.M.Johnst.) A.J.Paton A.Chev.
- (A.Chev.) A.J.Paton (Wight) A.J.Paton
- Coleus sylvestris
- Coleus gratus Coleus tetradenifolius
- Coleus amicorum
- Coleus cataractarum Coleus thalassoscopicus
- (P.I.Forst.) P.I.Forst. Coleus graveolens
- (B.J.Pollard) A.J.Paton (R.Br.) A.J.Paton (Perkins) A.J.Paton
- Coleus tomentifolius
- Coleus ater Coleus guerkei
- Coleus torrenticola (Briq.) A.J.Paton
- (P.I.Forst.) P.I.Forst. Coleus caudatus
- Coleus triangularis (A.J.Paton) A.J.Paton Coleus habrophyllus
- (S.T.Blake) P.I.Forst. & T.C.Wilson A.J.Paton (S.Moore) E.Downes & I.Darbysh.
- (P.I.Forst.) P.I.Forst. (A.J.Paton) A.J.Paton
- Coleus ulugurensis
- Coleus hadiensis Coleus unguentarius
- (Forssk.) A.J.Paton (Codd) A.J.Paton，异名：Plectranthus unguentarius
- Coleus caudatus Coleus velutinus
- (Trimen) A.J.Paton Coleus hairulii
- S.Moore Kiew, 2021 (van Jaarsv. & Hankey) A.J.Paton
- Coleus aegyptiacus
- Coleus auriglandulosus Coleus hallii
- Coleus venustus (J.K.Morton) A.J.Paton
- (P.I.Forst.) P.I.Forst. Coleus celsus
- Coleus verticillatus (Baker) A.J.Paton Coleus harmandii
- (A.J.Paton) A.J.Paton A.J.Paton
- (Doan ex Suddee & A.J.Paton) A.J.Paton (Guillaumet & A.Cornet) A.J.Paton & Phillipson
- Coleus villosus
- Coleus helferi Coleus wallamanensis
- (Hook.f.) A.J.Paton Coleus amiculatus
- Coleus centraliafricanus Coleus wugensis
- Gürke Coleus hereroensis
- A.J.Paton (Engl.) A.J.Paton (Codd) A.J.Paton
- Coleus xylopodus
- Coleus australis Coleus hijazensis
- Coleus yemenensis (Abdel Khalik) A.J.Paton
- A.J.Paton Coleus chevalieri
- Coleus zombensis (Baker) Mwany. (Forssk.) A.J.Paton
- (T.C.Wilson, P.I.Forst. & M.A.M.Renner) T.C.Wilson & P.I.Forst. (R.Br.) A.J.Paton Briq.
- Regel
- Coleus humulopsis (B.J.Pollard) A.J.Paton
- Coleus ciliatus Coleus hymalis
- (Bramley) A.J.Paton (J.R.I.Wood) A.J.Paton
- Coleus autranii
- Coleus idukkianus (J.Mathew, Yohannan & B.J.Conn) Smitha Coleus circinnatus
- Coleus igniarioides Briq.
- (Hedge) A.J.Paton & Phillipson (Ryding) A.J.Paton
- Coleus ignotus
- Coleus achankoviliensis Coleus amoenus
- Coleus clandulosus
- (Lukhoba & A.J.Paton) A.J.Paton
- Coleus inselbergii Coleus xerophilus
- Hook.f. (B.J.Pollard & A.J.Paton) A.J.Paton (T.C.Wilson & P.I.Forst.) T.C.Wilson & P.I.Forst.
- (Forssk.) A.J.Paton Coleus veyretiae
- Coleus barbatus
- Coleus insignis (P.I.Forst.) P.I.Forst. Coleus ventosus
- (Hook.f.) A.J.Paton Coleus venteri
- Coleus coeruleus
- Gürke Coleus trullatus
- Coleus insularis (P.I.Forst.) P.I.Forst. (Andrews) Benth. ex G.Don，异名：Plectranthus barbatus
- Gürke (P.I.Forst.) P.I.Forst.
- (Suddee) Suddee Coleus togoensis
- (A.J.Paton) A.J.Paton
- Coleus intraterraneus (Gürke) A.J.Paton & Phillipson Coleus suffruticosus
- (S.T.Blake) P.I.Forst. & T.C.Wilson (Gürke) A.J.Paton
- Coleus comosus
- Coleus stuhlmannii Coleus strobilifer
- Coleus kanyakumariensis (H.Keng) A.J.Paton Coleus splendidus
- Hochst. ex A.Rich., 1850，异名：''Plectranthus ornatus'' (sold as "''Coleus canina''" or "scaredy cat plant") (Shinoj & Sunojk.) Smitha
- (P.I.Forst.) P.I.Forst. Coleus splendens Coleus affinis
- Coleus bariensis
- Coleus kirkii (Radcl.-Sm.) A.J.Paton，异名：Plectranthus socotranus Coleus sigmoideus
- (Baker) A.J.Paton (A.J.Paton) A.J.Paton
- Coleus comosus Elmer
- Coleus scutellarioides Coleus kisanfuensis
- (Baker) A.J.Paton (Ryding) A.J.Paton Hochst. ex Gürke
- DeWild. Coleus schizophyllus Coleus scaber
- (Britten) A.J.Paton
- Coleus sallyae Coleus klossii
- (De Wild.) A.J.Paton
- (S.Moore) P.I.Forst. & T.C.Wilson Coleus anamallayensis
- Coleus confertiflorus Coleus rotundifolius
- (N.E.Br.) A.J.Paton Coleus kottalensis
- Coleus rhodesianum (Ryding) A.J.Paton (A.J.Paton) A.J.Paton
- A.Chev. Coleus rafidahiae
- (Buscal. & Muschl.) A.J.Paton Coleus pseudospeciosus
- Coleus batesii Coleus koulikoroensis
- Coleus psammophilus (T.J.Edwards & N.R.Crouch) A.J.Paton
- A.J.Paton Coleus porcatus
- Coleus congensis
- (Hook.f.) A.J.Paton Smitha & A.J.Paton (Gürke) A.J.Paton
- Bedd. (Baker) A.J.Paton (Gürke) A.J.Paton
- (Prain) A.J.Paton Coleus phulangkaensis
- (J.Mathew & B.J.Conn) A.J.Paton (Backer ex Adelb.) A.J.Paton
- Coleus petiolatissimus Coleus laetus
- (A.J.Paton) A.J.Paton Coleus penicillatus
- (P.I.Forst.) P.I.Forst.
- Coleus congestus Coleus parvifolius
- Gürke Coleus lageniocalyx
- Coleus pallidus (R.Br.) A.J.Paton
- Briq. Baker
- Coleus omissus
- Coleus batianoffii Coleus lanceolatus
- (Hiern) A.J.Paton (P.I.Forst.) P.I.Forst.
- (Bojer ex Benth.) A.J.Paton & Phillipson Coleus nigericus
- Coleus conglomeratus (Baker) A.J.Paton
- Coleus nepetifolius Coleus neochilus Coleus lancifolius
- (P.I.Forst.) P.I.Forst. (T.C.E.Fr.) Robyns & Lebrun
- (Bramley) A.J.Paton (Codd) A.J.Paton Coleus montanus
- (P.Beauv.) A.J.Paton
- Coleus lanuginosus
- (S.T.Blake) P.I.Forst. & T.C.Wilson Coleus minutus
- Coleus anamudianus
- Coleus crassus (J.K.Morton) A.J.Paton
- (Gürke) A.J.Paton Coleus melleri Coleus lateriticola
- (J.K.Morton) A.J.Paton (N.E.Br.) Culham
- (A.Chev.) Phillipson, O.Hooper & A.J.Paton Coleus marrubatus Coleus magnificus
- 马达加斯加鞘蕊花 Benth.
- Coleus bellus Coleus lateriticola
- Coleus lyratus
- A.Chev. Gürke Coleus livingstonei
- Coleus cremnus
- (P.I.Forst.) P.I.Forst. (Briq.) A.J.Paton Coleus latifolius
- (Smitha & Sunojk.) Smitha (P.I.Forst.) P.I.Forst. (B.J.Conn) A.J.Paton
- Hochst. ex Engl., 1892 (S.T.Blake) P.I.Forst. & T.C.Wilson
- Hochst. ex Engl., 1892
- Coleus kunstleri Coleus laxus
- (A.J.Paton) A.J.Paton Coleus huberi
- (T.C.Wilson & P.I.Forst.) T.C.Wilson & P.I.Forst. Coleus gossweileri
- Coleus cucullatus (Benth.) A.J.Paton
- Coleus fasciculatus Coleus leiperi
- Coleus dewildemanianus (A.J.Paton) A.J.Paton
- 杂交种：
- 归入本属的物种：
- Rabdosiella leemannii
- Solenostemon pubescens A.Chev.
- Saccostoma urticifolium N.Hahn Wall. ex Voigt
图库[编辑]
参考文献[编辑]
- ^ 1.0 1.1 1.2 Coleus Lour.. Plants of the World Online. Royal Botanic Gardens, Kew. [2020-07-28]. （原始内容存档于2020-10-27）.
- ^ 2.0 2.1 2.2 2.3 2.4 2.5 Paton, Alan J.; Mwanyambo, Montfort; Govaerts, Rafaël H.A.; Smitha, Kokkaraniyil; Suddee, Somran; Phillipson, Peter B.; Wilson, Trevor C.; Forster, Paul I. & Culham, Alastair. Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera. PhytoKeys. 2019, 129: 1–158. PMC 6717120 可免费查阅. PMID 31523157. doi:10.3897/phytokeys.129.34988.
- ^ Coleus Care. Gardening Know How. 2019-07-15 [2019-07-15]. （原始内容存档于2019-07-15）.
- ^ Morton, J.K. Plectranthus. Hedberg, I.; Kelbessa, E.; Edwards, S.; Demissew, S. & Persson, E. (编). Flora of Ethiopia and Eritrea 5. Addis Ababa: The National Herbarium, Addis Ababa University: 586–598. 2006. ISBN 978-91-971285-6-8., cited in Paton et al. 2019
- ^ Paton, A.; Mwanyambo, M. & Culham, A. Phylogenetic study of Plectranthus, Coleus and allies (Lamiaceae): Taxonomy, distribution and medicinal use (PDF). Botanical Journal of the Linnean Society. 2018, 188 (4): 355–376 [2020-12-22]. doi:10.1093/botlinnean/boy064. （原始内容存档 (PDF)于2020-11-30）.
- ^ Somran Suddee, Shuichiro Tagane, Phetlasy Souladeth, Deuantar Kongxaisavath, Sukid Rueangruea, Yoshihisa Suyama and Eizi Suzuki. 2020. Coleus bolavenensis (Lamiaceae), A New Species from Laos. Thai Forest Bulletin (Botany). 48(1), 82-85. DOI: 10.20531/tfb.2020.48.1.14
外部链接[编辑]
维基共享资源上的相关多媒体资源：Module:Commons_link第64行Lua错误：attempt to index field 'wikibase' (a nil value)
- EcoPort on P. rotundifolius (synonym of Coleus rotundifolius)
- PROTAbase on P. rotundifolius （页面存档备份，存于互联网档案馆） (synonym of Coleus rotundifolius)
- EcoPort on P. esculentus (with tuber photo) （页面存档备份，存于互联网档案馆） (synonym of Coleus esculentus)
- Lamiales Newsletter on P. esculentus （页面存档备份，存于互联网档案馆） (synonym of Coleus esculentus)
- Little-known African Tubers （页面存档备份，存于互联网档案馆） (discusses C. rotundifolius and C. esculentus)
Module:Taxonbar第160行Lua错误：attempt to index field 'wikibase' (a nil value)