涡虫纲
| 关于生物分类框涡虫纲 化石时期: 可能的奥陶纪记录[2]
| |
|---|---|
| 贝氏伪角扁虫(Pseudobiceros bedfordi) | |
| 贝氏伪角扁虫(Pseudobiceros bedfordi) | |
| 科学分类编辑 | |
| 界: | 动物界 Animalia |
| 演化支: | 肾管动物 Nephrozoa |
| 演化支: | 原口动物 Protostomia |
| 演化支: | 螺旋动物 Spiralia |
| 演化支: | 吮食动物 Rouphozoa |
| 门: | 扁形动物门 Platyhelminthes |
| 纲: | 涡虫纲 Turbellaria Ehrenberg, 1831 |
| 传统分类中包含的子类群 | |
| 传统分类中未包含,但在 演化层面上包含的演化支 | |
涡虫纲(学名:Turbellaria)是扁形动物门一个纲,传统上包括所有非寄生的扁形动物物种。大约有4500个物种,长度从1 mm(0.039英寸)到超过500 mm(20英寸)长的大型淡水物种[3]或陆生物种,如长达600 mm(24英寸)的柯氏广头地涡虫 (Bipalium kewense)。涡虫纲包含陆生及水生物种,大多数涡虫为海生,在底泥、岩石、海藻上爬行;淡水涡虫生活在池塘、溪流的石块下;陆生涡虫大多为夜行性,栖息在落叶或腐木等阴湿处。涡虫表具纤毛,摆动时积水呈涡状。所有较大型的生物都是扁平的,形状像带状或叶状,因为它们没有呼吸和循环系统,这意味着它们必须依靠扩散作用来进行代谢产物的内部运输。不过,许多小型生物的横截面是圆形的。它们大多是掠食者或食腐动物,都生活在水中或潮湿的陆地环境中。大多数的形态都是有性生殖,除了少数例外,所有的形态都是同时雌雄同体。
无腔动物(Acoelomorpha)和异涡虫属(Xenoturbella)属以前被归入涡虫纲,但现在已不再被视为扁形动物。所有专门寄生的扁形动物组成了单系的新皮总纲(Neodermata),人们一致认为它们是自由生活的扁形动物中一个小亚群的后裔。因此,传统定义中的 “涡虫纲 ”属于并系群。
分类[编辑]
涡虫纲传统上包括所有扁形动物门之下的非寄生物种。但根据分子生物学证据,涡虫类已被证实为并系群,原来属于寄生物种的三个纲(吸虫纲、单殖纲和绦虫纲)都是涡虫类物种的直系分支,加上涡虫纲内有两个目独立成另一门,较新的文献已不再使用本分类。
大多数一般人接触最多的涡虫类物种都属于三歧肠目。2008年以后的研究确认了:无肠目和纽涡虫目其实并不属于扁形动物,而是属于独立的无腔动物;而链涡虫目的物种亦较早与其他扁形动物门的物种分化。这两个分类群在较新的文献已不再被认为是涡虫类物种。然而在医学寄生虫学,基本上仍然沿用旧有的分类。
传统分类[编辑]
以下为传统的涡虫纲下属分类,可分为11个目:
- 无肠目 Acoela(现属无腔动物)
- 链涡虫目 Catenulida
- 单咽涡虫目 Haplopharyngida
- 卵黄皮目 Lecithoepitheliata
- 大口涡虫目 Macrostomida
- 纽涡虫目 Nemertodermatida(现属无腔动物)
- 多歧肠目 Polycladida
- 原卵黄目 Prolecithophora
- 原顺列目 Proseriata
- 棒肠目 Rhabdocoela
- 三歧肠目 Tricladida
演化树[编辑]
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涡虫纲(并系群) |
下属分类[编辑]
本纲包括以下目:[4]
- Proplicastomata
- Proporoplanidae
- 目的地位未定的科:
- Crassicolidae
- Dicteroidae
- Didymorchidae
- Diposthidae
- Discoprosthididae
- [[]]
- Euryleptididae
- Hoploplanidae
- Latocestidae
- Mucroplanidae
- Pericelidae
- Prosthiostomidae
- Stylochoididae
- Amyellidae
- Pterastricolidae
- Theamatidae
- Euryleptidae
- Anocellidae Haplopharyngidae
- [[]]
- Macrostomidae
- Palauidae
- Anonymidae Pleioplanidae
- [[]]
- Apidioplanidae
- Boniniidae
- Callioplanidae
- Candimboididae
- Catenulidae
- Cestoplanidae
- Chordariidae
- Chromoplanidae
- Cryptocelidae
- Didangiidae
- Diphosthiidae
- Discocelididae
- Ditremageniidae Euplanidae
- [[]] Gnesiocerotidae
- [[]] Koinocystidae
- [[]] Limnostylochidae
- [[]] Opisthogeniidae
- [[]] Planoceridae
- [[]] Retronectidae
- [[]] Tyrrheniellidae
- Polyposthiidae
- [[ Myoretronectes Noreña-Janssen & Faubel, 1996
- Paracatenula Sterrer & Rieger, 1974
- Retronectes Sterrer & Rieger, 1974
- Retronectes atypica Doe & Rieger, 1977
- Retronectes clio Sterrer & Rieger, 1974
- Retronectes euterpe Sterrer & Rieger, 1974
- Retronectes hyacinthe[5]
- Retronectes melpomene Sterrer & Rieger, 1974
- Retronectes sterreri Faubel, 1976
- Retronectes terpsichore Sterrer & Rieger, 1974
- Retronectes thalia Sterrer & Rieger, 1974]] Stylochocestidae
- Enantiidae
- Faubelidae
- Ilyplanidae
- Leptoplanidae
- Notoplanidae
- Phaenocelidae
- Pseudostylochidae
- Taenioplanidae
- [[ Theama Marcus, 1949
- Theama evelinae Marcus, 1949
- Theama forrestensis Bulnes & Faubel, 2003
- Theama mediterranea Curini-Galletti, Campus & Delogu, 2008[7]
- Theama occidua Sopott-Ehlers & Schmidt, 1975]] Stylochoplanidae
- [[ Cryptostylochus Faubel, 1983
- Distylochus Faubel, 1983
- Imogine Marcus & Marcus, 1968
- Kataria Faubel, 1983
- Katariana
- Leptostylochus Bock, 1925
- Mirostylochus Kato, 1937
- Pseudidioplana Prudhoe, 1989
- Stylochopsis Stimpson, 1857
- Stylochus Ehrenberg, 1831
- Stylochus alexandrinus Steinböck, 1937
- Stylochus aomori (Kato, 1937)
- Stylochus arenosus (Willey, 1897)
- Stylochus argus Czerniavsky, 1881
- Stylochus atentaculatus Hyman, 1953
- Stylochus bermudensis Verrill, 1902
- Stylochus californicus Hyman, 1953
- Stylochus castaneus Palombi, 1939
- Stylochus catus (Du Bois-Reymond Marcus, 1958)
- Stylochus ceylanicus (Laidlaw, 1904)
- Stylochus cinereus Willey, 1897
- Stylochus conglomeratus (Stimpson, 1857)
- Stylochus crassus Verrill, 1892
- Stylochus djiboutiensis Meixner, 1907
- Stylochus ellipticus (Girard, 1850)
- Stylochus exiguus Hyman, 1953
- Stylochus fafai (Marquina, Fernández-Álvarez & Noreña, 2015)
- Stylochus ferox Bock, 1925
- Stylochus flevensis Hofker, 1930
- Stylochus franciscanus Hyman, 1953
- Stylochus frontalis Verrill, 1893
- Stylochus (Distylochus) fundae Cuadrado, Rodriguez, Moro, Grande & Noreña, 2021
- Stylochus hamanensis Kato, 1944
- Stylochus hyalinus Bock, 1913
- Stylochus ijimai Yeri & Kaburaki, 1918
- Stylochus inimicus[6]
- Stylochus insolitus Hyman, 1953
- Stylochus (Distylochus) isifer (Du Bois-Reymond Marcus, 1955)
- Stylochus izuensis Kato, 1944
- Stylochus kimae (Jennings & Newman, 1996)
- Stylochus lateotentare (Lee, Beal & Johnston, 2005)
- Stylochus lesteri (Jennings & Newman, 1996)
- Stylochus limosus (Stimpson, 1857)
- Stylochus luteus (Muller J., 1854)
- Stylochus marmoreus Bock, 1925
- Stylochus (Distylochus) martae (Marcus, 1947)
- Stylochus matatasi Newman, Cannon & Govan, 1993
- Stylochus mcgrathi (Jennings & Newman, 1996)
- Stylochus mediterraneus Galleni, 1976
- Stylochus megalops (Schmarda, 1859)
- Stylochus meganae (Jennings & Newman, 1996)
- Stylochus meixneri Bock, 1925
- Stylochus melihertani (Bulnes, 2010)
- Stylochus meridianus Prudhoe, 1989
- Stylochus minimus Palombi, 1940
- Stylochus mistus Ramos-Sánchez, Carrasco-Rodríguez, Garcí-Madrigal & Bastida-Zavala, 2020
- Stylochus miyadii Kato, 1944
- Stylochus neapolitanus (Delle Chiaje, 1841)
- Stylochus nebulosus (Girard, 1853)
- Stylochus necopinata (Sluys, Faubel, Rajagopal & van der Velde, 2005)
- Stylochus oculifera (Girard, 1853)
- Stylochus orientalis Bock, 1913
- Stylochus pardalotus (Jennings & Newman, 1996)
- Stylochus pilidium (Goette, 1881)
- Stylochus pillidum (Goette, 1881)
- Stylochus plessisii Lang, 1884
- Stylochus pulcher Hyman, 1939
- Stylochus (Distylochus) pusillus (Bock, 1913)
- Stylochus pygmaeus Merory & Newman, 2005
- Stylochus qeshmensis (Maghsoudlou & Momtazi, 2014)
- Stylochus refertus Du Bois-Reymond Marcus, 1965
- Stylochus rutilis Yeri & Kaburaki, 1918
- Stylochus salis Cuadrado, Rodriguez, Moro, Grande & Noreña, 2021
- Stylochus salmoneus Meixner, 1907
- Stylochus sixteni (Marcus, 1947)
- Stylochus speciosus (Kato, 1937)
- Stylochus stellae (Marquina, Osca, Rodriguez, Fernandez-Despiau & Norena, 2014)
- Stylochus stellatus Jennings & Newman, 1996
- Stylochus suesensis Ehrenberg, 1831
- Stylochus tauricus Jacubowa, 1909
- Stylochus ticus Marcus, 1952
- Stylochus tripartitus Hyman, 1953
- Stylochus uniporus Kato, 1944
- Stylochus vesiculatus Jakubova, 1909
- Stylochus vigilax Laidlaw, 1904
- Stylochus zanzibaricus Laidlaw, 1903
- Stylochus zebra (Verrill, 1882)]] Stylochidae
- Stenostomidae
- Pseudocerotidae
- Plehniidae
- Laidlawiidae
- 目和科的地位未定的属:
- Diesing, 1862 Centrostomum
- Diesing, 1850 Conoceros
- Purasjoki, 1945 Darwinia
- Schmarda, 1859 Dicotylus
- Stimpson, 1857 Diporoplana
- Faubel, 1983 Ehrenberg, 1828
- Eumicrostomum Costa, 1882
- Extremodera Timoshkin & Zabrovskaya, 1985
- Geopaludicolia Le Conte, 1851
- Graffilla Verrill, 1893
- Kleinenbergia Stimpson, 1857 Girard, 1893
- Orthostoma Ehlers & Ehlers, 1981
- Platamonia Mamkaev, 1967
- Promesostum Örsted, 1843
- Thalattoplana Frölich, 1954
- Adenoperta
- Ehrenberg, 1828 Paradiscocelis
- Busch, 1851 Polyporus
- Antonius, 1968 Sphyrocephalus
- Bock, 1925 Typhlolepta
- Eukalyptorhynchia Diesing, 1850
- Aorchis Euvortex Pigulevsky, 1952
- Barker & Parsons, 1915 Komárek, 1920
- Graffia
- Itapis Graff, 1912
- Apoxenetes Oligocelis Ax, 1971
- Prudhoe, 1989
- Asoleina Proaphanostomum Hyman, 1959
- Schmarda, 1859
- Boemigia Sabussow, 1902
- Brabbinthes Allison, 1975
- Bubaloceras Matjašic, 1958
- Candimba Marcus, 1949
- Caspiodendrocoelum Dyganova, 1976
- Catenula Leblond, 1837
- Celidotis Diesing, 1862
- Cephalolepta Lang, 1884
- Coronhelmis Pereyaslawzewa, 1892
- Dicelis Grube, 1872
- Diplonchus Kohler, 1933
- Discoprpsthides Elasmodes
- Meixner, 1928 Eustomum
- Sekera, 1906 Gasterotrichula
- Diesing, 1861 Gnosonesimilia
- Levinsen, 1879 Homaloceraeus
- Marcus, 1952 Neocryptocelis
- Schmidt, 1852 Parapseudoceros
- Simonetta & Delle-Cave, 1978 Preminoma
- Vayssière, 1906 Tangarao
- Stimpson, 1857 Woodshollia
- Xaropoa Proantroporus
- Ørsted, 1843 Ehrenberg, 1831
- An-der-Lan, 1936 Busquet, 1899 Trachyplana
- Telostoma
- Korotneff, 1912 Tetracolylus Prudhoe, 1989
- Oligoclada Prudhoe, 1989
- Planoceroides Karling, 1967
- Procotylus Marcus, 1952
- Tetracotylus Graff, 1912
- Örsted, 1845 Rhodoplana
- Disorus Le Conte, 1851
- Euophthalmia Leidy, 1851
- Fridmaniella Graff, 1912
- Glossostoma Westblad, 1953
- Heterostylochus Schmarda, 1859
- Nautiloplana Pearse, 1938
- Palana Palombi, 1936
- Pocillophynchus Korotneff, 1908
- Smigrostoma Korotneff, 1912
- Tricelis
- Plehn, 1897 Brunet, 1973 Platydendron
- Kelaart, 1858 Penula
- Pararhynehella Faubel, 1983
- Orthostomum
- Stimpson, 1857 Neoplana
- Korotneff, 1912 Genneoceros
食物[编辑]
涡虫主要吃小的水生动物、动物尸体及动物内脏。寄生的吸收动物或植物体内的营养。
体型[编辑]
大部分涡虫的长度小于1cm,但热带的陆生种类长度可达120cm。
形状[编辑]
多为黑色,体前端有两个可感光的眼体,两侧对称。吻位于身体的三分之二处,且无肛门。有背、腹之分。
参见[编辑]
参考文献[编辑]
- ^ Dirk Knaust. Remarkably preserved benthic organisms and their traces from a Middle Triassic (Muschelkalk) mud flat. Lethaia. 2010, 43 (3): 344–356. doi:10.1111/j.1502-3931.2009.00196.x.
- ^ Dirk Knaust; André Desrochers. Exceptionally preserved soft-bodied assemblage in Ordovician carbonates of Anticosti Island, eastern Canada. Gondwana Research. 2019, 71: 117–128. Bibcode:2019GondR..71..117K. S2CID 134814852. doi:10.1016/j.gr.2019.01.016.
- ^ Ecology and Classification of North American Freshwater Invertebrates
- ^ Turbellaria. GBIF. [2023-10-09]. (原始内容存档于2023-02-18).
- ^ Niels W. L. Van Steenkiste, Alana Closs, Tyrel Froese & Brian S. Leander (2023) Molecular phylogenetic position of a rare and enigmatic meiofaunal flatworm from the Pacific Ocean: Retronectes hyacinthe sp. nov. (Platyhelminthes: Catenulida), Systematics and Biodiversity, 21:1, DOI: 10.1080/14772000.2023.2221236
- ^ Prof. Arturo Palombi (1931) Stylochus inimicus sp. nov. Policlade acotileo commensale di Ostrea virginica Gmelin delle Coste della Florida, Bollettino di zoologia, 2:1, 219-226, DOI: 10.1080/11250003109434873
- ^ M. Curini‐Galletti , P. Campus & V. Delogu (2008) Theama mediterranea sp. nov. (Platyhelminthes, Polycladida), the first interstitial polyclad from the Mediterranean, Italian Journal of Zoology, 75:1, 77-83, DOI: 10.1080/11250000701690525
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