勇蜗科
（重定向自Helicarionidae）
| 勇蜗科File:Infobox info icon2.svg | |
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| File:Helicarion cuvieri.jpg | |
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| File:Naturalis Biodiversity Center - RMNH.MOL.283157 1 - Fastosarion superba (Cox, 1871) - Helicarionidae - Mollusc shell.jpeg | |
| 科学分类 编辑 | |
| 界： | 动物界 Animalia |
| 门： | 软体动物门 Mollusca |
| 纲： | 腹足纲 Gastropoda |
| 目： | 柄眼目 Stylommatophora |
| 亚目： | 旋蜗牛亚目 Helicina |
| 下目： | 蛞蝓下目 Limacoidei |
| 总科： | 勇蜗总科 Helicarionoidea |
| 科： | 勇蜗科 Helicarionidae Bourguignat, 1877[1] |
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_-_Helicarionidae_-_Mollusc_shell.jpeg){kind=link}
勇蜗科（学名：Helicarionidae），又名鳖甲蜗牛科，鳖甲螺科[2]，是一个由陆生肺螺类半蛞蝓组成的科，皆为真有肺类柄眼类支序的成员，属于勇蜗总科（Helicarionoidea）[3]。
分布[编辑]
本科物种分布于古北界东部，马尔加什（马达加斯加、留尼汪及科摩罗）、印度、东南亚及澳大利亚[4]。
解剖[编辑]
本科的蜗牛都会在交配时伸出几丁质的交配器恋矢（love dart）插进对方的生殖孔，以交换精子[5]。
此外，本科物种的单倍体染色体介乎21到30之间[6]。
下级分类[编辑]
本科包括以下属：
- Hionella
- Iredale, 1941 曙螺属
- Uapuneutra Advena Gude, 1913
- Aenigmatoconcha C.Tumpeesuwan & S.Tumpeesuwan, 2017
- Amenixesta
- [[Stanisic, 2010
- Antiquarion boondah Hyman & Köhler, 2020
- Antiquarion chooreechillum Hyman & Köhler, 2020
- Antiquarion collinsae J.Stanisic, 2010
- Antiquarion evelynensis Hyman & Köhler, 2020
- Antiquarion limbus Hyman & Köhler, 2020
- Antiquarion occultus Hyman & Köhler, 2020
- Antiquarion ravenshoe Hyman & Köhler, 2020
- Antiquarion swausi Hyman & Köhler, 2020]] Apothapsia
- Austenia Robson, 1914
- Brevisentis Ehrenberg, 1831
- Cryptaustenia Dendronitor
- Stanisic, 2010 Fijia
- Hyman, 2007 Nesaecia
- Rhadella Hyman, Lamborena & Köhler, 2017
- Palmervillea Godwin-Austen, 1914
- Gude, 1913 Holkeion
- Gude, 1911 Stanisic, 2010
- Urazirochlamys Cucullarion Iredale, 1933
- Eufretum 吉拉蛞蝓属
- Godwin-Austen, 1908 Limpidarion
- Panulena Ancey, 1887
- 若通螺属 Vanmolia de
- Girasia Hyman & Köhler, 2020
- Nesarion Rotungia
- Winter, 2008 Hyman, 2007 Burnettia
- 疑难螺属 Cockerell, 1891 Stanisic, 1998
- Einasleighana H.B.Baker, 1941
- Gray, 1855 Howearion
- Baker, 1940 E.A.Smith, 1884
- Vercularion Saphtia de
- Iredale, 1933 Iredale, 1944
- Luinarion Parmavitrina
- Winter, 2008
- Dupontia Stanisic, 2010
- Gudeconcha 伊比蛞蝓属 Iredale, 1933
- Nesocyclus Satiella
- Baker, 1940 Parmella W.T.Blanford & Godwin-Austen, 1908
- Ibycus
- Baker, 1940 Attenborougharion
- G.Nevill, 1878 Belopygmeus
- Stanisic, 2010 Chalepotaxis
- Dancea Godwin-Austen, 1908
- Eurychlamys
- [[Heynemann, 1863
- 小玉蜗牛 Ibycus hiraseanus]] Malandena
- Parmellops Martens, 1860
- Iredale, 1944 Zagmena
- Gudeoconcha Iredale, 1933
- Nesonanina Sheaia
- Iredale, 1941 Zilch, 1960 小杜螺属
- Elatonitor Godwin-Austen, 1899 Iredale, 1944
- Insulorbis C.R.Boettger, 1918
- Iredale, 1937
- Sigaloeista Baker, 1940
- Mauka Periclocystis
- Shea & Griffiths, 2010 Buffetia
- [[Ancey, 1887
- Neutra 扁楯蛞蝓属 Sitala
- Petalochlamys H.Adams, 1865 Baker, 1940
- Irenella Baker, 1940
- Godwin-Austen, 1907，叶楯蛞蝓属 Dedrolamellaria
- [[W.T.Blanford, 1863
- Durgella aborense Godwin-Austen, 1916
- Durgella assamica Gondwin-Austen, 1881
- Durgella birmanica (Pfeiffer, 1847)
- Durgella christianae (Theobald, 1864)
- Durgella concinna Blanford & Godwin-Austen, 1908
- Durgella densestriata Vermeulen, T.-S.Liew & Schilthuizen, 2015
- Durgella dwipana (Gude, 1903)
- Durgella edeana Godwin-Austen, 1910
- Durgella erratica (Godwin-Austen, 1888)
- Durgella hosei Godwin-Austen, 1891
- Durgella khasiaca Godwin-Austen, 1883
- Durgella kumahensis (Theobald & Stoliczka, 1872)
- Durgella levicula (W.H.Benson, 1859)
- Durgella libas Solem, 1966
- Durgella mairangensis Godwin-Austen, 1898
- Durgella naharaniensis Godwin-Austen, 1914
- Durgella nulla Pholyotha & Panha, 2024[9]
- Durgella pentata Pholyotha & Panha, 2024[9]
- Durgella rhaphiellus (E.von Martens, 1867)
- Durgella rogersi Godwin-Austen, 1907
- Durgella russeola (Morelet, 1865)
- Durgella salius (W.H.Benson, 1859)
- Durgella seposita (W.H.Benson, 1859)
- Durgella siamensis Möllendorff, 1902
- Durgella sundana B.Rensch, 1930]] Expocystis
- Gude, 1913 Medyla
- Pitcairnia H.B.Baker, 1941
- Baker, 1940 Iredale, 1941
- Amenixesta fifensis Iredale, 1941
- Amenixesta hunsteini (E.A.Smith, 1887)
- Antiquarion D.T.Holyoak & G.A.Holyoak, 2020
- Hyman & Köhler, 2017 Belopygmaeus
- Baker, 1938 Iredale, 1945 Caldwellia
- Ancey, 1904
- Ellarion Iredale, 1937
- Germain, 1918 Kalidos
- Gude, 1911 Preston, 1913
- Sodaleta 勇蜗属
- Gude, 1911 Minororbis
- Plegma Iredale, 1937
- Durgellina Iredale, 1941
- Helicarion Baker, 1940
- Nukupiena Sophina
- Pravonitor Benson, 1859 Férussac, 1821
- Kermarion Baker, 1940
- Iredale, 1937
- H.Adams, 1873 Colmanarion
- Delinitesta Thiele, 1928
- Fastosarion Iredale, 1944
- Mistarion Pseudaustenia
- Hyman & Ponder, 2010 Iredale, 1939 Cockerell, 1891
- Westracystis Waihoua
- Hiloaa Iredale, 1941
- Oafatua Tarocystis
- Habe, 1946 Ubiquitarion
- Baker, 1940
- Iredale, 1933 Epiglypta
- Iredale, 1933 Baker, 1940 Kipua
- Baker, 1940 Baker, 1925
- Pachystyla Iredale, 1937
- Gude, 1911 Stanisic, 2010 Germain, 1918
- Teraia
- Rhyssotopsis Torrecystis
- Baker, 1940 Montanocystis
- 奥氏蛞蝓属 Bathia Solem, 1989
- Burmochlamys
- Stanisic, 2010 Ctenophila
- Echonitor Pilsbry, 1893
- Hiona Stanisic, 2010
- Opara Thularion
- Sitalarion
- Solem, 1966 Iredale, 1937 Stanisicarion
- Pseudosaphtia de Stanisic, 1993 W.T.Blanford, 1863
- Sivella Cooke, 1940 Laconia
- Baker, 1940 Winter, 2008
- 昔塔螺属 Hyman, 2007
- Sesara
- Godwin-Austen, 1918 Dendrolamellaria Iredale, 1937
- Fenestrarion
- Gray, 1855 Muangnua 圆鳖甲蜗牛属
- Pseudosesara Stanisic, 2010
- Laidlaw, 1932 Rasama Baker, 1940
- Mörch, 1852 H.Adams, 1867 Solem, 1962
- Quirosella
- Hionarion Solem, 1966
- Ovachlamys Tropicystis
- Pseudophasis Pseudoguppya Pholyotha & Panha, 2022
- Caracolla 隐奥氏蛞蝓属 Ancey, 1882
- Preston, 1913 Erepta
- Stanisic, 2010 Baker, 1940 Laocaia
- Habe, 1946 Clench, 1958
- Pittoconcha
- Iredale, 1937 Peloparion
- Uafatua
- Parmacochlea
- Kuzminykh, 1999 Mysticarion
- Rapafila Baker, 1940 Nitor
- Martens, 1860 Levidens
- Baker, 1940
- Eddiella Albers, 1850 Harmogenanina
The following two subfamilies were recognized in the taxonomy of Bouchet & Rocroi (2005):[3]
- 亚科 Helicarioninae Bourguignat, 1877 - synonyms: Pseudotrochatellinae A. J. Wagner, 1905;[10] Ereptinae Godwin-Austen, 1908;[11] Xestinae Gude & B. B. Woodward, 1921;[12] Sesarinae Thiele, 1931;[13] Nitoridae Iredale, 1937;[14] Epiglyptidae Iredale, 1944;[15] Gudeoconchidae Iredale, 1944[15]
- 亚科 Durgellinae Godwin-Austen, 1888[16]
属[编辑]
下列本科所有的属：
Helicarioninae亚科[编辑]
- Epiglypta
- Gudeoconcha Pseudotrochatella
- [[Iredale, 1914[3]]] G. Nevill, 1881[3] Erepta
- Sesara Albers, 1860[3]
- Helicarion Pilsbry, 1893[3]
- [[Albers, 1850[3]]] Férussac, 1821 - type genus of the family Helicarionidae[3] Preston, 1913 [21]
- Xesta
- Nitor Albers, 1850[3]
- Wilhelminaia Gude, 1911[3]
Durgellinae亚科[编辑]
- 族 Durgellini
- Durgella
- Sitala H. Adams, 1865[3]
- Satiella Blanford, 1863 - type genus of the subfamily Durgellinae[3]
- [[Godwin-Austen, 1908[3]]] Benson, 1859[3] Sophina
- 族 Girasiini
亚科未定[编辑]
- Advena Buffetia
- Mathewsoconcha
- Mysticarion
- Nancibella
- 圆鳖甲蜗牛属 Ovachlamys
- Habe, 1946， 卵楯蛞蝓属 Diastole
- Panulena
- Pittoconcha
- Quintalia
- Dolapex
- Ryssota
- Thapsia
- Trichotoxon
- Zingis Elisolimax
- Atrichotoxon
- Harmogenanina
- Iredaleoconcha
- 恰里螺属 Kaliella
- Lutilodix
- Philonesia
亲缘支序图[编辑]
以下的系统发生学分支图显示了根据Hausdorf (2000)所描述蛞蝓下目各科之间的亲缘关系[4]：
| 蛞蝓下目 Limacoidei |
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参考文献[编辑]
- ^ Bourguignat J. R. (1877). "Description de deux nouveaux genres algériens, suive d'une classification des families et des genres de Mollusques terrestres et fluviatiles du système européen". Bulletin de la Société des Sciences Physiques et Naturelles de Toulouse 3(1): 49-101. page 64.
- ^ 施乃普. 綠島小形貝殼. 中国贝志 (中华民国贝类学会). 1975, 2: 33–46 [2018-08-28] （中文（繁体））.
- ^ 3.00 3.01 3.02 3.03 3.04 3.05 3.06 3.07 3.08 3.09 3.10 3.11 3.12 3.13 3.14 Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. Classification and nomenclator of gastropod families. Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 978-3-925919-72-5. OCLC 61351079. OL 42989175M. S2CID 83847814.
- ^ 4.0 4.1 Hausdorf, B. Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal. Journal of Biogeography. 2000, 27 (2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x （English）.
- ^ 記用蝸牛來作實驗. 2013-07-19 （中文（繁体））.
- ^ G. M. Barker. The Biology of Terrestrial Molluscs. CABI. 2001: 139. ISBN 978-0-85199-318-8.
- ^ Chanidaporn Tumpeesuwan and Sakboworn Tumpeesuwan. 2017. Discovery of An Overlooked Helicarionid Land Snail (Helicarionidae: Durgellinae) from northeastern Thailand, with Description of A New Genus and New Species, and Note on Radula Morphology and Genital System. RAFFLES BULLETIN OF ZOOLOGY. 65; 181–188.
- ^ Arthit Pholyotha, Chirasak Sutcharit, Piyoros Tongkerd and Somsak Panha. 2021. Systematic Revision of the Limestone Karst-restricted Land Snail Genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with Description of A New Species. European Journal of Taxonomy. 767(1), 55-82. DOI: 10.5852/ejt.2021.767.1487
- ^ 9.0 9.1 Arthit Pholyotha, Chirasak Sutcharit, Ngwe Lwin and Panha Somsak. 2024. Unveiling the Diversity of the Semislug Gnus Durgella Blanford, 1863 (Eupulmonata: Helicarionidae) from Thailand and Myanmar, with Description of Two New Species. Zool Stud. 63:14. DOI: 10.6620/ZS.2024.63-14.
- ^ Wagner A. J. (1905). Denkschriften der Mathematisch-Naturwissenschaftlichen Klasse der Kaiserlichen Akademie der Wissenschaften 77: 365.
- ^ Godwin-Austen H. H. (1908). "On the Animals of Genera and Species of Mascarene Land-Mollusca belonging to the family Zonitidae, collected by Monsieur E. Dupont". Annals and Magazine of Natural History (8)2: 422-436. 432. Plate 9-11.
- ^ Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 185.
- ^ Thiele (1931). Handbuch der systematischen Weichtierkunde 1(2): 620.
- ^ Iredale T. (1937). The Australian Zoologist 9(1): 2.
- ^ 15.0 15.1 Iredale T. (1944). The Australian Zoologist 10(3): page 326 and page 328.
- ^ Godwin-Austen H. H. (1888). Land and freshwater Mollusca of India 1(6): 253.
- ^ Godwin-Austen H. H. (1900). In: Sykes Mollusca. In: Sharp (ed.) Fauna Hawaiiensis 2(4): 283.
- ^ Blanford W. T. & Godwin-Austen H. H. (1908). The fauna of British India. Mollusca. Testacellidae and Zonitidae, page 283.
- ^ Schileyko (2003). Treatise on Recent terrestrial pulmonate molluscs, part 10: 1319.
- ^ Collinge (1902). The Journal of Malacology 9(3): 71, 73.
- ^ Ho, Wai Hoong. A Review Of The Land-Snail Fauna of Singapore. The Raffles Bulletin of Zoology. 1995, 43 (1): 91–113 [2017-02-16]. （原始内容存档于2017-02-16） （English）.
外部链接[编辑]
- File:Wikispecies-logo.svg 维基物种上的相关信息：勇蜗科
- 勇蜗科. 美国国家生物技术信息中心（NCBI）. 424154. 已忽略未知参数
|taxon=(帮助) - Video of a Helicarionid in Sulawesi （页面存档备份，存于互联网档案馆）
