端足類
| 端足目File:Infobox info icon2.svg 化石時期:
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|---|---|
| File:Gammarus roeselii.jpg | |
| Gammarus roeseli | |
| 科學分類 編輯 | |
| 界: | 動物界 Animalia |
| 門: | 節肢動物門 Arthropoda |
| 亞門: | 甲殼亞門 Crustacea |
| 綱: | 軟甲綱 Malacostraca |
| 亞綱: | 真軟甲亞綱 Eumalacostraca |
| 總目: | 囊蝦總目 Peracarida |
| 目: | 端足目 Amphipoda Latreille, 1816[1] |
| 亞目 | |
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| Module:Wd第196行Lua錯誤:attempt to call field 'getGlobalSiteId' (a nil value) | |
端足類(學名:Amphipoda),亦作片腳類,是一個沒有甲殼及兩側扁平的目級甲殼類動物。其學名的意思是「不同的足」,是指它們擁有不同的附屬肢形態。它們長1-340毫米不等,棲息在所有的水生環境及一些潮濕的陸生環境。當中有7000個物種,其中5500個被分類到鉤蝦亞目中,其餘的則分到餘下的二或三個亞目。
特徵[編輯]
端足類的身體共分13節,可以分類為頭部、胸部及腹部。[2]頭部與胸部融合,有兩對觸角及一對複眼。[3]頭上亦有一個口器,不過大部份的口器都是隱藏的。[4]胸部及腹部各有不同,且有不類別的腳。它們一般都是兩側扁平的,且沒有甲殼。[3]胸部有八對單肢附足,第一對是輔助口器的,其後四對向前,最後三對向後。[3]鰓位於胸節,循環系統是開放的,擁有心臟,血青蛋白在血淋巴中運送氧至身體組織。觸角上有特別的腺來控制吸收及排出鹽份。[2]腹部分為兩部份:前尾有游泳肢,後尾有尾節及三對腹足,腹足並不會組成尾扇。[3]
端足類一般短於10毫米,但最長活生生的端足類可以達28厘米,是生活在太平洋5300米水深處的地方。[5]於大西洋發現的標本重組後就發現長達34厘米,屬於Alicella gigantea的。[6]最細小的就只有1毫米長。[7]端足類的大小受到溶氧供應的限制,如在海拔3800米的的的喀喀湖中,端足類就可以生長達22毫米;而在海拔455米的貝加爾湖,其中的端足類就長達90毫米。[8]
繁殖及生命周期[編輯]
成熟的雌性有育幼袋,可以帶著受精卵[2],直至幼體孵化為止。[3]雌性的年紀越大就會產更多的卵。卵的死亡率約為25–50%。[2]它們沒有幼體階段,卵會直接孵化出幼生。經過6次脫殼後,它們就達至性成熟。[2]一些物種在脫殼後會吃下自己的殼。[2]
多樣性及分類[編輯]
卡爾·林奈在其第十版的《自然系統》(Systema Naturae)中描述了兩種端足類,不過描述得十分簡單且適用於差不多所有端足類。[9]現時約已描述了7000種端足類,分類在3或4個亞目中。鉤蝦亞目下就包含了多過5500種,包括了所有淡水及陸生物種。[4]英高蟲亞目包含了2科約40種[10],但這類有時被認為是與鉤蝦亞目相同,而非獨立的一個亞目。[4]
端足類因其體型細小而很難辨認,往往都需要將之解剖才能確認。生態及環境研究很多都會將它們混為一談。[11]端足類的分類尚未得到共識,當中以鉤蝦亞目內的關係產生最多混亂。[12]以下是其中一個建議的分類法:[12]
- 鉤蝦亞目(Gammaridea)
- Ampeliscoidea
- Crangonyctoidea
- Dexaminoidea
- Eusiroidea
- Gammaroidea
- Hadzioidea
- Iphimedioidea
- Kurioidea
- Leucothoidea
- Liljborgioidea
- 琴鈎蝦總科 Lysianassoidea
- Melphidippoidea
- Oedicerotoidea
- Pardaliscoidea
- Phoxocephaloidea
- Stegocephaloidea
- Stenothoidea
- Synopioidea
- Talitroidea
- Thurstonelloidea
- 麥桿蟲亞目(Caprellidea)
- Caprellida
- Caprelloidea
- Phtisicoidea
- Cyamida
- 鯨虱科(Cyamidae)
- 𧊕亞目(Hyperiidea)
- 體箱𧊕次目(Physosomata)
- 頭箱𧊕次目(Physocephalata)
- 端足目包括以下科:
- Acanthogammaridae
- Acanthonotozomellidae Ambasiidae
- Caspicolidae
- Paracalliopiidae Falklandellidae
- [[]]
- [[]]
- Paracrangonyctidae Aristiidae
- Gamaridae Paragammaropsidae
- [[]] Cebocaridae
- Paraleptamphopidae
- Gammaracanthidae
- [[]]
- [[]] Paramelitidae
- Gammarellidae
- Paramphithoidae
- Ceinidae 鈎蝦科
- Paraphronimidae Gammaridae
- [[]]
- [[]] Aetiopedesidae
- Artesiidae
- Gammaroporeiidae Pardaliscidae
- [[]] Cheidae
- Perthiidae
- Giniphargidae
- [[]]
- [[]] Photidae
- 強螯鈎蝦科 Hadziidae
- Phoxocephalidae Ampeliscidae
- Cheirocratidae
- Phoxocephalopsidae Haustoriidae
- [[]]
- [[]]
- Phronimidae Astyridae
- Hirondelleidae Phrosinidae
- [[]] Cheluridae
- Platyischnopidae
- [[]]
- [[]]
- [[]] Pleustidae
- Hyalellidae
- Plioplateidae Hyalella S.I.Smith, 1874
- Hyallela troglofugia Bastos-Pereira, Oliveira & Ferreira, 2018
- Hyalella yashmara Tomikawa, Kawasaki, Leiva & Arroyo, 2023[13]
- Chevaliidae
- Plioplateiidae Hyalidae
- [[]]
- [[]]
- Atylidae
- Hyperiidae Podosiridae
- [[]] Chillagoeidae
- Pontogammaridae
- Hyperiopsidae
- [[]]
- [[]] Pontoporeiidae
- Iciliidae
- Priscillinidae Amphilochidae
- Chiltoniidae
- Priscomilitaridae Ingolfiellidae
- [[]]
- [[]]
- Pronoidae Australomicroprotopidae
- Ipanemidae Proscinidae
- [[]] Chuneolidae
- Protodulichiidae
- Iphigenellidae
- [[]]
- [[]] Pseudamphilochidae
- Iphimediidae
- Pseudingolfiellidae
- Colomastigidae
- Pseudocrangonyctidae Isaeidae
- [[]]
- Acanthonotozomatidae
- Alicellidae Austroniphargidae
- Ischyroceridae
- Regaliidae
- Condukiidae Salentinellidae
- Iulopididae
- [[]]
- [[]]
- Sandroidae Izinkalidae
- Scinidae
- Amphithyridae Conicostomatidae
- Scopelocheiridae
- Kairosidae
- [[]]
- Seborgiidae Baikalogammaridae
- Kamakidae
- Sensonatoridae
- Corophiidae Sicafodiidae
- [[]]
- [[]]
- [[]]
- Sophrosynidae Kergueleniolidae
- Stegocephalidae
- [[]] Crangonyctidae
- Stenothoidae
- Kotumsaridae
- [[]]
- 藻鈎蝦科 Bateidae
- Kuriidae
- Synopiidae
- Crangoweckeliidae Talitridae
- Lafystiidae
- [[]]
- [[]]
- Thamneidae Lanceolidae
- Acidostomatidae
- Ampithoidae Cressidae
- Thurstonellidae
- Laphystiopsidae
- [[]]
- Tryphanidae Bathyporeiidae
- Lepechinellidae
- Tryphantidae
- Crymostygidae Tryphosidae
- Lepidepecreellidae
- [[]]
- [[]]
- Typhlogammaridae Lestrigonidae
- Uhlorchestiidae
- [[]] Crypturopodidae
- Unciolidae
- Leucothoidae
- [[]]
- Allocrangonyctidae Behningiellidae
- Liljeborgiidae
- Uronyctidae
- Curiotalitridae Urothoidae
- Luciobliviidae
- [[]]
- [[]]
- Valettiidae 鯨虱科 Lycaeidae
- Valettiopsidae
- Anapronoidae Cyamidae
- Vemanidae
- Lycaeopsidae
- [[]]
- 琴鈎蝦科 Vicmusiidae Bogidiellidae
- Lysianassidae
- Vitjazianidae
- Cyclocaridae
- [[]]
- [[]]
- [[]]
- Zobrachoidae
- Maeridae
- Cyllopodidae Magnovidae
- [[]]
- 異鈎蝦科 Bolttsiidae
- Makawidae Cyphocarididae
- Maxillipiidae
- [[]]
- Megalanceolidae
- Anisogammaridae Cyproideidae
- Megaluropidae
- [[]] Bougisidae
- Melitidae
- Cystisomatidae Melphidippidae
- [[]]
- [[]]
- Mesogammaridae
- Dairellidae Metacrangonyctidae
- [[]]
- Amaryllididae Brachyscelidae
- Metaingolfiellidae
- Derjugianidae Metalanceolidae
- [[]]
- [[]] Microphasmidae
- Aoridae
- Dexaminidae Microprotopidae
- [[]]
- Brevitalitridae Microprotopodidae
- Didymocheliidae
- [[]]
- [[]]
- Micruropidae
- Dikwidae
- Micruropodidae
- [[]] Calliopiidae
- Mimonecteolidae
- Dogielinotidae Mimonectidae
- [[]]
- [[]] Mimoscinidae
- Adeliellidae
- Archaeoscinidae 麥稈蟲科 Dulichiidae
- Miramarassidae
- [[]] Caprellidae
- Miramassidae
- Dussartiellidae Najnidae
- [[]]
- [[]]
- Neomegamphopidae
- Endevouridae
- Neoniphargidae Zaramillidae
- [[]]
- Wandinidae Amathillopsidae
- Caprellinoididae
- Nihotungidae Vibiliidae
- [[]]
- Eophliantidae
- Valettidae Niphargidae
- [[]]
- Urohaustoriidae Uristidae
- [[]]
- Tulearidae Nuuanuidae
- [[]]
- Arcitalitridae
- Epimeriidae Trischizostomatidae
- [[ Chevreuxiella Stephensen, 1915
- Chevreuxiopsis Halfter & Coleman, 2019
- Chevreuxiopsis franki Halfter & Coleman, 2019[14]
- Danaella Stephensen, 1925
- Parachevreuxiella Andres, 1987
- Thoriella Stephensen, 1915]] Ochlesidae
- Thoriellidae
- Chevreuxiopsis Halfter & Coleman, 2019
- Temnophliantidae
- [[]] Stilipedidae
- Caprogammaridae Odiidae
- Sternophysingidae
- Sinurothoidae
- Eriopisidae
- [[]]
- [[]]
- Sebidae
- [[]]
- Sanchoidae Ommatogammaridae
- [[]] Rakiroidae
- [[]]
- Eulimnogammaridae Pseudoniphargidae
- [[]] Opisidae
- Protorchestiidae
- [[]]
- Prolanceolidae
- Carangoliopsidae Otagiidae
- [[]]
- Pontogeneiidae
- Eupronoidae
- Podoprionidae Podoceridae Oxycephalidae
- [[]]
- [[]] Platyscelidae
- [[]]
- [[]] Pachyschesidae
- Phreatogammaridae
- Argissidae
- Eurytheneidae
- Phliantidae Pagetinidae
- [[]]
- Parascelidae Parargissidae
- [[]]
- Carinogammaridae Pakynidae
- Paraliscidae
- [[]] Paracercopidae
- Eusiridae
- [[]] Pallaseidae
- [[]]
- [[]]
- [[]]
- Oedicerotidae Parabogidiellidae
- Microscinidae
- Macrohectopidae
- Exoedicerotidae
- Kergueleniidae 壯角鈎蝦科 Paracalliopidae
- Hornelliidae
- 科的地位未定的屬:
- Anisopus Austrohyalella
- Mesocyamus Margolis, McDonald & Bousfield, 2000
- Mesohyalella Bousfield, 1996 Bousfield, 1996
- Moanamoera Staude, 1995
- Paraneohela
- Oldevig, 1959 Axelboekia
- Poeckilogammarus
- Templeton, 1837
- Praegmelina Derjavin, 1929
- Rhithromoera
- Staude, 1995 Cylindrimolaris
- Rosagammarus McMenamin, Zapata & Hussey, 2013
- Stenia Ortiz & Lalana, 1999 Dana, 1849
- Transitroides
- Variogammarus Tachteew, 1995 Electicus
- Lowry & Stoddart, 1997 Apocyamus
- Gainella Chevreux, 1911
- Gammaroidorum Margolis, McDonald & Bousfield, 2000 Jarzembowski, Chény, Fang & Wang, 2020
- Humilomoera Staude, 1995
- 細螯蝦屬 Leptochela
- Boeck, 1876 Bousfield & Poinar, 1994 Polichinellobizarrocomic
化石紀錄[編輯]
估計端足類源自三疊紀,但其化石紀錄很稀有,至今只有11個物種標本被發現,最遠追溯至上始新世。[15]
生態[編輯]
端足類分佈在所有水生環境,由淡水至含有兩倍鹽份的海水也有。[2]它們是水生生態系統的重要組件。[11]大部份鉤蝦亞目物種都是淺水底棲的。𧊕亞目是浮游的。[3]很多都是與凝膠狀動物,如水母、管水母目、放射蟲及櫛水母等具有共生關係,而大部份𧊕亞目的生命周期都會涉及這些凝膠狀動物。[16]
一些擊鉤蝦科是陸生的,棲息在潮濕的環境,如葉堆等。[17]它們的分佈很廣,包括屬於以往岡瓦那大陸部份,與及部份歐洲及北美洲地區。
約有750種端足類是穴居的,分佈在差不多所有適合的環境。它們的多樣性中心在地中海盆地、北美洲東南部及加勒比地區。[18]
與其他甲殼類(如等足目)比較,端足類較少是寄生的。最著名寄生的端足類是鯨虱,它們是腹背扁平的,有強壯及很大的爪,用來將自己附在鬚鯨之上。它們是唯一寄生的甲殼類不懂游泳的。[19]
大部份端足類都是吃碎屑或食腐動物[2],另有一些是吃藻類、雜食性或掠食細小昆蟲及甲殼類。[2][3]它們會用最前兩對有爪的腳來抓食物吃。[2]
參考[編輯]
- ↑ 引用錯誤:無效的
<ref>標籤,未定義名稱為ITIS_93294的參考文獻內容文字。 - ↑ 2.00 2.01 2.02 2.03 2.04 2.05 2.06 2.07 2.08 2.09 Sam Wade, Tracy Corbin & Linda-Marie McDowell. Class Crustacea. Critter Catalogue. A guide to the aquatic invertebrates of South Australian inland waters (PDF). Waterwatch South Australia. 2004. ISBN 1-876562-67-6. (原始內容 (PDF)存檔於2009-10-17).
- ↑ 3.0 3.1 3.2 3.3 3.4 3.5 3.6 Order Amphipoda. Guide to the marine zooplankton of south eastern Australia. Tasmanian Aquaculture & Fisheries Institute. 2008. (原始內容存檔於2008-07-20).
- ↑ 4.0 4.1 4.2 John R. Holsinger. What are amphipods?. Old Dominion University. [2010-04-07]. (原始內容存檔於2011-07-20).
- ↑ J. Laurens Barnard, Darl E. Bowers & Eugene C. Haderlie. Amphipoda: The Amphipods and Allies. Robert H. Morris, Robert Hugh Morris, Donald Putnam Abbott & Eugene Clinton Haderlie (編). Intertidal Invertebrates of California. Stanford University Press. 1980: 559–66. ISBN 0-8047-1045-7.
- ↑ J. Laurens Barnard & Camilla L. Ingram. The supergiant amphipod Alicella gigantea Chevreux from the North Pacific Gyre. Journal of Crustacean Biology. 1986, 6 (4): 825–39 [2010-06-04]. (原始內容存檔於2020-07-31).
- ↑ T. Wolff. The fauna of Rennell and Bellona, Solomon Islands. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 1969, 255 (800): 321–43 [2010-06-04]. (原始內容存檔於2020-06-10).
- ↑ L. S. Peck & G. Chapelle. Reduced oxygen at high altitude limits maximum size. Proceedings of the Royal Society B. 2003, 270: S166–7. doi:10.1098/rsbl.2003.0054.
- ↑ S. Pinkster. Redescription of Gammarus pulex (Linnaeus, 1758) based on neotype material (Amphipoda). Crustaceana. 1970, 18 (2): 177–186 [2010-06-04]. (原始內容存檔於2020-05-03).
- ↑ R. Vonk & F. R. Schram. Ingolfiellidea (Crustacea, Malacostraca, Amphipoda): a phylogenetic and biogeographic analysis. Contributions to Zoology. 2003, 72 (1): 39–72 [2010-06-04]. (原始內容存檔於2013-12-03).
- ↑ 11.0 11.1 J. K. Lowry & R. T. Springthorpe. Introduction. Amphipoda: Families. Australian Museum. [2010-04-05]. (原始內容存檔於2006-02-21).
- ↑ 12.0 12.1 Joel W. Martin & George E. Davis. An Updated Classification of the Recent Crustacea (PDF). Natural History Museum of Los Angeles County. 2001: 132 [2010-06-04]. (原始內容存檔 (PDF)於2013-05-12).
- ↑ Ko Tomikawa, Yoshimi Kawasaki, Alfonso Miranda Leiva and Nilton Deza Arroyo. 2023. Description of A New thermal Species of the Genus Hyalella from Peru with Molecular Phylogeny of the Family Hyalellidae (Crustacea, Amphipoda). Invertebrate Systematics. 37(4); 254-270. DOI: 10.1071/IS22060
- ↑ Svenja Halfter; Charles Oliver Coleman; Chevreuxiopsis franki gen. n., sp. n. (Crustacea, Amphipoda, Thoriellidae) from the deep sea southwest of Tasmania. 2019
- ↑ E. L. Bousfield & G. O. Poinar, Jr. A new terrestrial amphipod from tertiary amber deposits of Chiapas province, Southern Mexico. Historical Biology. 1994, 7 (2): 105–14. doi:10.1080/10292389409380448.
- ↑ G. R. Harbison, D. C. Biggs & L. P. Madin. The associations of Amphipoda Hyperiidea with gelatinous zooplankton. II. Associations with Cnidaria, Cteuophora and Radiolaria. Deep-Sea Research. 1977, 24: 465–88. doi:10.1016/0146-6291(77)90484-2.
- ↑ M. A. Minor & A. W. Robertson. Amphipoda. Guide to New Zealand Soil Invertebrates. Massey University. 2010-03-05 [2010-04-07]. (原始內容存檔於2010-05-10).
- ↑ Horton H. Hobbs, III. Crustacea. John Gunn (編). Encyclopedia of Caves and Karst Science (PDF). Routledge. 2003: 254–7 [2010-06-04]. ISBN 978-1-57958-399-6. (原始內容存檔 (PDF)於2004-12-28).
- ↑ Tim Goater. Parasitic Amphipoda. Interactive Parasitology. Vancouver Island University. 1996-05-04 [2010-04-07]. (原始內容存檔於2010-07-14).
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